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601 Cards in this Set

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what is biology?
this is the scientific study of living things
definition of living things?
these are all the organisms descended from a single-celled ancestor
characteristics of living organisms
1. consist of one or more cells
2. contain genetic information
3. use genetic information to reproduce themselves
4. are genetically related and have evolved
5. can convert molecules from their environment into new biological molecules
6. can extract energy from teh environment and use it to do bilogical work
7. can regulate their internal environment
living systems evolve through...
these things evolve through differential survival and reproduction
what is the central theme of biology?
evolution - what to biology?
who said and when?
"nothing in biology makes sense except in the light of evolution"
Theodosius Dobzhansky
1973 - what did he write
what did Charles Darwin propose?
that all living organisms are descended from common ancestors - evolution by natural selection
what was charles darwin interested in?
geology and natural science
when and what did Darwin do on his voyage around the world
he accepted a position on the HMS Beagle for a 5-year survey voyage- most important in galapagos islands
what did darwin study in the glapagos islands
he observed that species were similar to, but not the same as, species on the mainland of south america

he also realized that species varied from island to island
what did Darwin postulate from his studies on the galapagos island
he postulated that species had reached the islands from the mainland, but then had undergone different changes on different islands
what did Darwin propose as an explanatory theory for evolutionary change
1. species change over time
2. divergent species share a common ancestor
3. the process that produces the change is natural selection
what helped darwin realize natural selection was possible
humans select for desired traits when breeding animals (artificial selection) (foxhound vs beagle) then it is possible that nature to do the same selection process
Columba livia
300 varieties of dove derived form teh wild rock dove
Darwin raised these as a hobby
pigeons
darwin postulated that natural selection could occur through...
what occurs though differential selection and reproductive success
define differential survival?
not every organism will have the same chance for survival
when years of research from darwin
1831- 1858
who wrote letter to darwin proposing an explanation of natural selection almost identical to darwin's
alfed russel wallace
who wrote a paper with darwin - when and who submitted
July 1 1858
alfred russel wallace
linnean society of london

- work not understood - not received well
what book did darwin publish and when

- what did this book do?
origin of species in 1859
- exhaustive evidence from many different fields to support evolution and natural selection
- even incluced geological evidence
who were darwin adn wallus influenced by?
economist Thomas Malthus - who had published An Essay on the Principle of Population
what did Thomas Malthus publish and when
An Essay on the Principle of Population
1838
- populations of all species have the potential for rapid increase
- but this does not occur in nature, so death rate must also be high
if populations increase exponetially, supplies increast arithmatically, who dies if this is not seen in nature?
very young and very old
natural selection - define
differential contribution of offspring to the next generation by various genetic types of belonging to the same population
in evolutionary biology adaptation refers to :
1. the process by which characteristics appear to be useful evolve
2. a phenotypic characteristic that makes it more likely the organism will survive
traits that increase the probability of survival and successful repordiction will increase in populations - give examples
fall - leaves prevent moisture loss by losing leaves
carnicouous plants - adapted to living in poor soild areas - need amino acids not available in soil, so plant consumes flys
The basic structural and physiological unit of all living organisms is the
cell is the basic...
All living organisms acquire _______ from their environment.
energy
Which of the following represents a correct ordering of the levels of complexity at which life is studied, from most simple to most complex?
Cell, tissue, organ, organism, population, community
A prerequisite for the survival of life on land was the accumulation of
O2 how was this important to suvival on land?
A species is...
a group of organisms that look alike and are capable of interbreeding. - what is this?
Although Darwin’s voyage aboard the HMS Beagle took him around the world, most of his observations on natural history were made on the continent of _______ and its nearby islands.
South America

Galapagos Islands

- who studied here?
Darwin was influenced by _______ in his recognition that populations of animal and plant species have the potential to expand rapidly, but that in nature, this population growth is usually checked by high death rates.
Thomas Malthus
Domain Eukarya includes
plants, amimals and fungi
Heterotrophs obtain their energy from
other organisms
Name one thing that is not a feature of scientific hypotheses?
They are unable to be falsified.
define living thing
all the diverse organism descended from a single-celled ancestor that evolved almost 4 billion years ago
modern cell theory
- all cells come from preexisting cells
- all cells are similar in chemical composition
- most of the chemical reactions of life occur in aqueous solution within cells
- complete sets of genetic information are replicated and pased on during cell division
-viruses lack cellular stucture but remain dependent on cellular organisms
evolution
change in the genetic makeup of biological populations through time
adaptations
structural, physiological or behavioral traits that enhance an organisms chances of survival and reporduction in its environment
sexual selection
selection due to mate choice
genetic drift
the random fluctuation of gene frequenceies in a population due to chance events
cells genome
the sum total of all the DNA molecules it contains
DNA
doxyribonucleic acid

long sequences of four different subunits called nucleotides
necleotides
DNA base pairs
Genes
specific segments of DNA encoding the information the cell uses to make proteins
proteins
molecules govern the chemical reactions within the cells and form much of the organisms structure
nutrients
supply the organism with energy and raw materials for carrying out biochemical reactions
snythesis
building new complex molecules and atructures form smaller chemical units
metabolism or metabolic rate
the sum total of all the chemical transformations and other work done in all the cells of an organism
differentiated
specialized
assemblages of differentiated cells are organized into...
tissues
different tissue types are organized to form ... that accomplish specific functions
organs
organs whose functions are interrelated can ge brouped into
organ systems
(ex digestive system)
fuctions of cells, tissues, organs, and organ systems are all integral to the multicellular...
organism
a group of individuals of the same species that interact with one another is a
population
populations of all species that live and interact in teh same area are called a
community
communities together with their abiotic environment contstitute an
ecosystem
ecosystems exchange energy and create earth's...
biosphere
prokaryotes
first unicellular organism consisting of DNA and other biochemicals enclosed in a membrane (bacteria)
photosyntesis
changed the nature of life on earth - build up of O2 and allowed life to move on land
eukaryotes
have nuclei and other internal components
binomial
biologists give each species a distinctive scientific name formed from two latinized names
first - genus
second - species
genus
a group of species that share a cecent common ancestor
systematists
scientists who study the evolution and classification fo lifes diverse organisms
phylogentic trees
document and diagram evolutionary relationships as part of an overarching tree of life
name three major domains
archaea
bacteria
eukarya
domains of single celled prokaryotes
archaea adn bacteria
donmain of cells whose mitrochondria and chloroplasts may have originated from the ingestion of prokaryotic cells
eukarya
name three major groups of multicellular eukaryotes
- what group did they evolve from?
plants, fungi, and animals

- each devolved from a different group of the eukaryotes generally referred to as protists
scientific method
1. make observations
2. speculate, ask a question
3. form a hypothesis to answer the question
4. make a prediction: what else wold be true if your hypothesis is correct?
5. design and conduct an experiment that uses quantifiable data to test your prediction
hypothesis
tentative answers to questions
name two types of experiments
controlled and comparitive

controlled - one variable changes
comparitive - can't controll all variables - simply gathering and comparing data from different sample groups
null hypothesis
assuming there is no difference
what was charlse darwin passionately interested in?
geology and natural science
what happened in 1831 with Darwin?
he was recommended for a position on teh HMS Beagle for a 5 year voyage around the world
what did darwin observe with respect to parents and their offspring?
he observed that, though offspring tended to resemble their parents, they are not identical
what animal did darwin breed?
pigeons
what animal did darwin study on the galapagos island?
finches - specifically their beaks
artificial selection
people breeding with specific characteristics in mind
do individuals evolve?
no - populations evolve
who evolves?
populations, not people
population
a group of individuals of the same species that live and interbreed in a particular geographic area
adaptation
the processes by which useful characteristics evolve; and the characteristics themselves
those in a population who do not adapt to the environment...
do not survive and do not pass on their genes
gene
segment of DNA that codes for a trait
allele
different form of a gene
phenotype
the physical expression of an organisms genes

ex brown eyes, the ability to digest gluten, predisposition to cancer
genotype
the allele combination of a trait

Bb
homozygous
both alleles are the same

BB or bb
heterozygous
alleles are different

Bb
brassica oleracea
through artificial selection 6 different plants can come from this single species depending on what trait is bred for:
1. cabbage - terminal buds
2. cauliflower - flower clusters
3. brussels sprouts - lateral buds
4. broccoli - stems and flowers
5. kohlabi - stem
6. kale - leaves
drosophila
fruit fly
mendelian populations
locally interbreeding groups
allele frequencies
p = their proportion in the gene pool

estimated by counting alleles in a sample of individuals
how are allele frequencies estimated?
by counting alleles in a sample of individuals
equation for allele frequency
p = # copies of the allele in population / sum of alleles in the population
poymorphic
more than one morph at that locus - ex Aa
equation
frequency of allele A =
2NAA + NAa / 2N

N= total number of individuals in the population
frequency of allele a =
2Naa + NAa / 2N

N= total number of individuals in the population
If p is frequency of allele A
and q is freq of allele a
then p+q =
1

with hardy weinberg equilibrium taken into consideration
if there is only one allele at a locus, its frequency is
1 and the population is monomorphic at that locus
monomorphic
one morph at that locus - AA only or aa only
if certain conditions are met - teh genetic structure of a population...
does not change over time
hardy-weinberg equilibrium
describes a model situation in which allele frequencies do not change

not ideal - to many variable in real life
- mutations, unequal offspring
conditions that must be met to maintain hardy-weinberg equilibrium
1. mating is random
2. populationsize is infinate. large populations aren't affected by genetic drift
3. no gene flow- no migration into or out of the population
4. no mutation
5. natural selection does not affect survival of any genotypes
if conditions hold, after one generation, genotype frequencies occur in these proportions
AA p squared
Aa 2pq
aa q spared
hardy-weinberg equilibrium is what type of hypothesis?
null hypothesis - assumes evlolutionary forces are absent
known evloutionary mechanisms that make hardy-weinberg non-realistic
mutation
gene flow
genetic drift
nonrandom mating
natural selection
genetic drift
results form random changes in allele frequencies

allele frequencies drift away from the original frequencies

effects small populations that colonie a new region
-colonizing population is unlikely to have all the alleles present in the whole population
in large populations, genetic drift can influence frequencies of alleles that dont affect....
survival and reproduction
population bottleneck
genetic drift can reduce the genetic drift - occurs in populations that are reduced to a snall number of individuals
founders effect
equivalent to a bottleneck
natual selection
adaptation occurs when some individuals in a population contribute more offspring to the next generation.

allele frequencies change in a way that adapts individuals to the environment that influenced that reproductive success.
natural selection acts on...
phenotype
fitness
reproductive contribution of a phenotype (lifespan and #offspring) to subsequent generations
changes to (what) of different phenotypes in a population leads to change in allele frequencies
relative success
natural selection can act on charaters with quantitative variation in three ways
1. stabilizing selection
2. directional selection
3. disruptive selection
stabalizing selection
preserves average phenotype
directional selection
favors individuals that vary in one direction
disruptive selection
favors individuals that vary in opposite directions from the average

- birds with average bills died out due to lack of access to food
sexual selection
special type of natural selection, which acts on characters that determine reproductive success

if an individual survives but does not reproduce, it makes no contribution to the next generation

favors traits that increase the chances of reproduction
intrasexual selection
traits that may improve ability to compete for mates - ex bright colors, long horns
intersexual seletion
be more attractive to the opposite sex

reiably demonstrate the fitness of the bearer to the choosing sex
humans have also influenced the evolution of other species
insecticiseds, antibiotics, hunting, moving species (plants, animals), changing climate
species
a "kind" of organism - group of actually or potentially interbreeding natual populations, which are reproductively isolated from other such groups

historically - fixed and unchanging creations of God - based on morphological characteristics
problems with concept of species
1. all dogs come from canis lupus
2. red wolf - hybrid or species, do we protect it?
3. asexually reproducing species
4. ring species (salamanders)
geographic speciation
populations are separated by a physical barrier

- once barrier is removed populations mingle but do not interbreed
allopatric speciation
Allopatric speciation (from the ancient Greek allos, "other" + Greek patra, "fatherland") or geographic speciation is speciation that occurs when biological populations of the same species become isolated due to geographical changes such as mountain building or social changes such as emigration. The isolated populations then undergo genotypic and/or phenotypic divergence as: (a) they become subjected to different selective pressures, (b) they independently undergo genetic drift, and (c) different mutations arise in the populations' gene pools.
sympatric speciation
does not require physical isolation

disruptive selection required

- ex birds with average beaks die off due to lack of access to food

most commonly occurs by polyploidy
polyploidy
duplicaiton of the whole set of chromosomes
autopolyploidy
chromosome duplication in a single species
allopolyploidy
combining of chromosomes from two species
reproductive isolation
1. prezygotic
a. habitat
b. temporal
c. behavioral
d. mechanical
e. gametic
2. postzygotic
a. low hybrid viability
b. hybrid infertility
prezygotic
before fertilization barriers prevent fertilization
habitat isolation
individuals live in different habitats
temporal isolation
individuals breed at different times (frogs)
behavioral isolation
individuals fail to recognize each other as potential mating partners
mechanical isolation
individuals cannot copulate with each other because of anatomical differences in their genitalia
gametic isolation
sprem cannot fertilize eggs
postzgotic barriers
prevent the production of viable fertile offspring
low hybrid viability
hybrid offspring are less likely to survive, either as zygotes (dying during development) or as adults (dying before they can mate)
hybrid infertility
hybrids may survive, but they cannot produce offsprng (a mule is the sterile offspring of a horse and a donkey)
Which of the following modes of selection leads to a reduction in variation but no change in the mean?
Stabilizing selection
The binomial system of nomenclature, by which we still classify species, was originated by
Carolus Linnaeus
The biological species concept is most closely associated with which scientist
Ernst Mayr
Geographical speciation is also known as
allopatric speciation
Sympatric speciation requires
disruptive selection.
Despite inhabiting overlapping ranges, the western spotted skunk and the eastern spotted skunk do not interbreed. One reason for this lack of interbreeding is that the western species breeds in early fall and the eastern species breeds in late winter to early spring. This is an example of _______ isolation.
temporal
when and who recommended darwin for 5 year trip on the HMS Beagle
John Henslow
1831
after his ship voyage what did Darwin propose
1. species change over time
2. diverent species share a common ancestor
3. mecahnism for change is natural selection
who did similar research to Darwin and published an presented work together with Darwin
Alfred Russel Wallace
1858
who was Wallace and Darwin influenced by (an essay on the principle of population)?
Thomas Malthus (1838 wrote essay)
what was called when Gregor Mendal's publications were rediscovered and people studied the mechnistic aspects of evolution as well as the broad evolutionary patters evident in nature?
population genetics
modern synthesis
heritable trait
characteristic at least partly determined by the organisms genes
population genetics three goals?
1. explain the pattrens and organization of genetic variation
2. explain the origin oand maintenance of genetic variation
3. understand the mechanisms that cause changes in teh allele frequencies in populatinos
gene pool
sum of all copies of all alleles at all loci found in a population
what can demonstrate the existence of considerable genetic variation in populations?
lab experiments
ex. fruit fly - high and low bristle count bred from selection of regular bristle fly generations
mendelian populations
locally interbreeding groups
frequency
an allele's proportion in the gene pool at a particular locus
AA, Aa, aa - what is it called in a population when more than one genotype found
polymorphic
AA or aa - what is it called in a population when only one genotype found
monomorphic and allele is said to be fixed
genetic structure
frequencies on different alleles at each locus and the frequencies of different genotypes in a mendelian poluation describe that poluations what?
who deduced the conditionss that must prevail if the genetic structure of a population is to remain the same over time?
and when?
Godfrey Hardy Wilhelm Weinberg

Hardy-Weinberg equilibrium
1908

only applies to sexually reproducing organisms
hardy-wienberg equilibrium

after one generation of random mating - what are the genotype frequencies?
AA - p squared
Aa - 2pq
aa - q squared
if generation has more homozygous individuals and fewer heterozygous individuals that would be expected this condition is called?
heterozygote deficiency
why is hardy-weinberg equilibrium important if populations in nature never meet the conditions?
1. equation is useful for predicting the approximate genotype frequencies of a population from its allele frequencies
2. model describes the conditions required for there to be no evloution in a population
how does calculating allele frequencies allow us to measure evolutionary change?
allele frequencies measure the amount of genetic variation in a population and comparing this over time allows for chnges (or lack there of) can be considered over time (evolution)
principle of natural selection?
two meanings of adaptation?
differential survival and reproductive success

1. processes by which characteristics that appear to be useful to their bearers evolve (evolutionary mechanisms that produce them) and the characteristics themeselves

2. phenotype characteristic that has made it more likely for an organism to survive and reproduce
evolutionary mechanisms that make hardy-weinberg equilibrium unrealistic
1. mutation
2. gene flow
3. genetic drift
4. nonrandom mating
5. selection
mutation
any change in the nucleotide sequences of an organisms DNA
explain study of genetic composition of founding populations
washington and chili vs europe with fruit flys
europe - 80 different chromosomal inversions
washington, chili - 20 different inversions

only a small part of the total genetic variation found in europe reached americas
give example of directional selection
horns of cattle - longer horned parents could defend their young better and so they were able to pass on genes
give example of stabalizing selection
baby birth weight
babies with higher and lower birth weights die at a higher rate than babies born at average weights
when and what book did darwin write about sexual selection
1871
The sescent of Man, and selection in relation to sex

these features (that would inhibit survival) either improved the ability of their berers to compete for access to mates (intrasexual - ex brighter color makes it easier for mate to see them) or made their bearers more attractive to memebers of the opposite sex (intersexual)
name two experiments that test sexual selection
1. widowbirds - tail length
2. zebra finches - carotenoid levels, immune sytems
give example of poulation bottleneck
greater prairie chickens - reduced to about 50 in 1990s
what are the constriants of evolution
lack of genetic variation can prevent evolution of potentiall favorable traits
- if not in population, it can't be passed on

humans have also influenced
-hunting, environment, pesticides, antibiotics
who developed the biological species concept and when
Ernst Mayr in 1940

species are groups of actually or potentially interbreeding natual populations, which are reproductively isolated from other such groups
who developed the binomial nomenclature and what called (two parts)
Carolus Linnaeus (Genus species)
problem with species concept
hybrids or species (ex red wolf), asexual species, ring species (slamanders),
what is required in sympatic speciation and what is not required
disruptive selection required
physical isolation
sympatic speciation most commonly occurs by
polyploidy
who developed the binomial system of biological nomenclature by which species are named today?
carlolus linnaeus
morphological species concept
construct that assumes a species consists of individuals that "look alike" and that individuals that don't look alike belong to different species
cryptic species
instances in which two or more morphologically indistinguishable species do not interbreed (salamanders)
lineage species concept
species as branches on the tree of life
reproductive isolation
a state in which two populations can no longer exchange genes (ex salamanders)
who proposed the biological species concept and what is it?
Ernst Mayr
"species are groups of actually or potentially interbreeding natural populations which are reproductively isolated from other such groups"
allopolyploidy
from teh combining of the chromosomes of two different species
reinforcement
strengthening of prezygotic barriers
layers of rock
strata
who first discovered strata and when?
Nicolaus Steno
17th century
How do scientists date ancient events?
by ratio of isotope C14 : C12

current during lifetime, C14 no longer uptakes after death

can go back up to 50,000 years

carbon dating
when were eras established
before ages of rocks were known
how do scientists date events? how is the history of life divided?
divided into geologic eras, which are subdivided into periods
bounderies are bosed on
changes in fossils
when did level of oxygen start to increase
after first photo-synthetic eukaryotes - (after first eukayotes)

about 1.5 billion years ago
organisms are classified according to
the period they were discovered
who established the binomial nomeclature system and when
1707-1778
carolus linnaeus

Genus species
ex Homo sapiens
hierarchical system of nomenclature devloped by Linnaeus
Domain
Kingdom
Phylum
Class
Order
Family
Genus
Species

King Phyllup Came Over For Good Spaghetti
Phylogentic tree
shows evolutionary relationships among lineages
describ in 5 steps how phylogenetic tree is created
1. a species, population, or gene at one point in time
2. becomes a leneage as we follow its decendants through time
3. a split occurs when the ancestral lineage divides into two decendant lineages
4. and each leneage continues to evolve independently as different traits arise (red dots)
5. the lineages continue to split and a phylogenetic tree emerges
phylogenetic tree

define root
the common ancestor for the group on the left
phylogenetic tree

define node
the splits in branches - indicate a division of one lineage into two
what do teh positions of the nodes on the time scale indicate
indicate the times of the corresponding speciation events
how can branches be changed and does this affect the tree?
branches can be rotated around any node without changing the meaning of the tree
synapormorphies
shared derived traits define a branch of a phylogenetic tree

note - trait at node is where species diverge
phylogenetic tree

define ingroup
group of organisms of primary interest
phylogenetic tree

define outgroup
closely related species known to be outside the group of interest
taxonomists use taxa for what?
use taxa for biological classification
classification

tree - wgat is a complete branch on the tree called
clade
paraphyletic group
group includes the common ancestor and some, but not all, of the ancestor's decendants (pink)
polyphyletic group
does not include the common ancestor of the group (yellow)
monophyletic group
includes the common ancestor and all decendants of that ancestor (blue)

monophyletic group can be removed from tree with a single cut
Domains - list
Archaea
Bacteria
Eukarya
name 4 approaches to classifying organisms
1. two-kingdome system - Linnaeus
a. Plantae
b. Animalia
2. five-kingdom system - Whittaker
a. Monera
b. Protista
c. Fungi
d. Plantae
e. Animalia
3. six-kingdom system - Woese
a. Eu-bacteria
b. Archae-bacteria
c. Protista
d. Fungi
e. Plantae
f. Animalia
4. three-domain system - Woese
a. Bacteria
b. Archaea
c. Eukarya
Archaea - describe
once celled organisms
live in extreme environments (bad water - death vally, hot springs, etc)
Protists - describe
one celled eukaryotic
example - amoeba
Fungi - describe
excrete enzymes and absorb food
example - mushrooms
what organisms believed to be first
prokaryotes
origin of mitrochondria and chloroplasts by
endosymbiotic - taking in without destroying - the two organisms then work together
endosymbiotic
taking in without destroying - the two organisms then work together
which is beleived to occur first - endosymbiotic origin of mitrochondria or endosymbiotic origin of chloroplasts?

when did this happen on domain tree?
endosymbiotic origin of mitrochondria

happened after split from archaea
members of all domains can?
1. conduct glycolosis
2. replicate DNA conservatively
3. have DNA that encodes peptides
4. produce peptides by transcription and translation using the same genetic code
5. have plasma membranes and ribosomes
example of archaea
halophiles
methanogens
halophile
example of archaea

"salt lover" - live in high salt and up to ph 11.5
methanogens
example of archaea

produce methane
thermoplasma
example of archaea

no cell wall, themophilic and acidophilic - lives in coal deposits
sulfobus
example of archaea

live in hot sulfur springs 70-75*C, pH 2-3
acidophilic hyperthermophiles
example of archaea

maintains interal temp of 7.0
name three bacteria shapes
helical
bacilli (rod)
cocci (spherical)
to group bacteria
shape
gram staining (yes.no)
nutrition (parasites, photoautotrophs)
metabolism
High GC, low GC
what kind of bacteria changed the earths atmosphere
cynobacteria
"pond scum"

forms colonies that have three cell types - spores, vegetative cells and heterocysts
what was the environement doing when eukaryotic cells were developed?
changing from anaerobic to aerobic
major events that occorred in evolution of eukaryotic cells are still...
conjectural (based on guesswork)
endosymbiosis - who developed theory
Lynn Margulis
eukaryotes that are neither plants, animals, or fungi are...
protists - what are these?

also called mirobial eukaryotes (though not all are microbial)
how can protists or microbial eukaryotes be classified?
1. by locomotion
2. motil - cilia or flagella
3. amoeboid motion - pseudopods
4. immotile- stationary
5. reproduction - most are asexual, but some are sexual
characteristics of heterotrophic fungi
1. absorptive heterotrophy (nutrition)
2. saprobes
3. parasites
4. mutualists


6 major groups
absorptive heterotrophy (nutrition)
secrete digestive enzymes and absorb nutrients
saprobes
absorb sutrients from dead organic matter
parasites
abosrb nutrients from living hosts
mutualists
livein intimate association with another organism that benefits both. Mutually beneficial
name 6 groups of fungi
1. microsporidia
2. chytrids
3. zygomycota
4. glomeromycota
5. ascomycota
6. basidiomycota
dashed line on phylogenetic tree means?
it indicates the probable paraphyly of the group
budding

ex. yeast
mitosis followed by asymmetrical cell division
fungi reproduce rapidly when?
nutrient supplies dwindle

ex - after it rains
predatory fungi do what?
they trap microscopic organisms
lichen
fungus and photosynthentic mircroorganism

mutalism
30,000 species
what are the major groups of eukaryotes
kingdoms:
1. Plantae
2. Animalia
3. fungi
4. Protista
all chloroplasts have what in common
all trace back to a single incidence of endosymobiosis
plantae phylogenetic tree has how many clades?
several
which is the closest living relative of the land plant?
charophytes
ex. coleochaeta alge

stonewarts (genus Chara)
sister group to plants
classification of land plants
nonvascular land plants
ex. moss - not tall, very green, found in moist environments

vascular plants (includes seed plants)
*****************************

what are key characteristics that evolved and allowed plants to live on land

******************************
1. waxy cuticle
2. Gametangia
3. protected embryos
4. absence of herbivores (initially)
5. vascular tissue
waxy cuticle
prevents dehydration
gametangia
cases to enclose gametes
vascular tissue
xylem - conducts water and minerals from soil upwards
phloem - conducts photosythetic products to roots for storage

allows plants to grow taller and large root systems support their height
nonvascular plants
do not have xylem and phloem - do not grow tall

need to live near water to reproduce

lack typical leaves, roots, and stems
what is considered oldest evolutionary plant
liverwort
root - common ancestor of seed plants - what are the nodes?
what is the outgroup and what is the ingroup?
gymnosperms (naked seeds) are the ingroup

angiosperms (flowering plants) is the outgroup
what makes an animal and animal?
eukaryotic
multicellular
heterotrophic metabolism
movement (exception coral)
if single cellular and eukaryotic
protist
to classify all animals we have to be...
general

1. body plan is the general structure
2. symmetry of body
3. body cavity structure (# layers)
4. segmentation of body
5. external appendages that move body
animal feeding strategies
1. filter feeders
2. herbivores
3. predators
4. parasites
5. detritivore (eats dead organic material)
sessile
stationary
motile
moving
what is a deuterosome?
characterized by three early developmental patterns:
1. radial cleavage
2. mouth forms opposite the blastopore
3. coelom develops from mesodermal pockets that bud off from the cavity of the gastrula
evidence from DNA sequencing supports the monophyly of the deuterostomes:
1. echinoderms
2. hemichordates
3. chordates

there are fewer species of deuterostomes than protostomes
living fossil
coelacanth
coelacanth - why still here?
no natural predeters
able to reproduce
isolated
have multiple food source
not commerical game fish
difference between amphibians and reptiles
1. amphibians existed before reptiles
2. amphibian reproduction tied to water (vernal pools)
3. can live on land (amphibians must stay moist)
4. external fertilization (amphibians) and internal fertilization (reptiles)
ex of amphibians and reptiles
Reptiles include alligators, crocodiles, turtles, and snakes. Amphibians include salamanders, toads, and frogs.
reptiles evolved before or after amphibians?
after
what adaptations did reptiles go through to survive on the land from water?
1. better lungs
2. hard shelled eggs
3. internal fertilization
4. scaly skin
5. kidneys excreted concentrated urine to prevent water loss
human evolution
primates
opposable digits
binocular vision
bipedal (two legs)
prehesile tail
tail can grasp
neutral allele
allele that does not affect the fitness of an organism
phylogeny
evolutionary history of relationships
node
a point at which lineages diverged in the past
taxon (taxa)
group of species that we designate or name
clade
any taxon that consists of all the evolutionary descendants of a common ancestor
systematics
the study and classification of biodiversity
homologous
any features shared by two or more species that have been inherited from a common ancestor
ancestral trait
a trait that was already present in teh ancestor of a group
derived trait
a trait found in a descendent that differs from its ancestral form
synapomorphies
derived traits that are shared among a group of organisms (viewed as evidence of the common ancestry of the group)
convergent evolution
independently evolved traits subjected to similar selection pressures may become superficially similar

(bones in bats and birds homologous, but wings are not - evolved independently)
evolutionary reversal
a character may revert from a derived state back to an ancestral state
homoplasies
similar traits generated by convergent evolution and evolutionary reversals
morphological species concept
a construct that assumes a species consists of individuals that look alike
lineage species concept
species as branches on the tree of life - each species has a history that starts at a speciation event (one lineage on teh tre splits into two) and ends either at extinction or another speciation event, at which time the species produces two daughter species
reproductive isolation
a state in which two polulations can not longer exchange genes
polyploidy
the duplication of sets of chromosomes within individuals
allopolyploidy
the combining of the chromosomes of two different species
autopolyploidy
chromosome duplication in a single species
sedimentary rocks
rocks formed by the accummulation of grains on teh bottom of bodies of water
strata
the oldest layers fo rock
precambrian era
more than 3 billion years
life consisted of microscopic prodaryotes; eukaryotes eveolved about 2/3 of the way through the era
all the plants living at a particular time or place
flora
all the animals living at a particular place or time
fauna
cambrian explosion
a geologically rapid diversification of life - took millions of years - oxygen concentration was approaching current level
name three eras
plaeozoic, mesozoic, cenozoic
synapormorhies
shared derived traits define a branch of a phylogenetic tree
taxonomists
use taxa for biological clasification
classifying organization - 4 version
1. LInneaus 2 kingdom
2. Whittaker - 5 kingdom
3. Woese - 6 kingdom
4. Woese - 3 domain
all domains
going
round
past
every
primate makes ripples

conducts glycolosis
replicate DNA
produce proteins by translation and transpcription
encodes proteins by using DNA
plasmind membranes and ribosomes
bacteria - how classified
shape
gramp pos/neg
nutrition
metabolism
archaea - how classified
unique environments
sulfobus
methangens
eukaryotes - what kingdoms
protists
animilia
fungi
plantae
protists - how classified
locomotion
- flagellum, cillia
- amoeboid
- immotile
reproduction
- asexual or sexual
biofilms
microbials (prokaryotes) put down a gel like sticky polysaccaride matirx that makes it hard to remove organism
what makes an animal and animal
every
mouse
has
its
momement

eukaryote
mulitcellular
heterotroph
internal digestion
movement (ex coral -sperm)
fungi - how classifed (heterothrpic)
1. absorptive heterotrophy
2. saprobes
3. parasites
4. mutualists
abosroptive heterotrophy
secrete digestive enzymes and absorb nutrients
saprobes
abosorb nutrients from dead organic matter
parasites
absorb nutrients from living hosts
protosomes
mouth first
- blastophore becomes mouth
deuterostomes
rear first
- blastophore becomes anus
detritivore
eats dead or organic matter
sessile
stationary
prokaryotes have distintive cell walls
peptidoglycan - gram pos/neg
photoautotrophs
photoheterotrophs
photoautotrophs - make own food (only need sunlight)

photoheterotroph - make own food, but require some compounds made from other organisms
protists
eukaryotes that are not plant, animal, fungi
sporophyte
spore plant
gametophyte
gamete plant
heteromorphic
alternation of generations, where generations alternate morphology
isomorphic
generations do not alternate
diatom
symetrical protist
plantae - name 5 major clades
1. glaucophytes
2. red algae
3. chlorophytes
4. land plants
5. charophytes
gametangia
gametes form wihtin specialized sex organs
what are unicellular fingi know as
yeasts
radial cleavage
fertilized egg dividing in an even pattern
what makes a mamal?
1. sweat glands
2. mamary glands
3. hair
4. four chambered heart
angiosperms - what are they
flowering plants - vasular plants with seeds enclosed in modified leaves called carpels (seeds enclosed during pollination)

more recent evolution
two clades of angiosperms
97% fall into these:
monocots

eudicots
monocots
narrow-leaved; grasses, lilies, orchids, palms
eudicots
broad-leaved; soybeans, roses, sunflowers, maples
what does remaining 3% of angiosperms look like?
structure similar to eudicots:
ex - water lilies, magnoliids
cotyledons
A leaf of the embryo of a seed plant, which upon germination either remains in the seed or emerges, enlarges, and becomes green
how many cotyledons does eudocots/monocots have?
monocots - one
eudocots - two
characteristics of monocots
1. one cotyledons
2. parallel veins in leaves
3. flowers usually in multiples of three
4. primary vascular bundles in stem are scattered
characteristics of eudicots
1. two cotyledons
2. veins in leaves are usually netlike
3. flower parts usually arranged in fours or fives
4. arrangement of primary vasular bundles in stem are in a ring
carpels
modified leaves of angiosperms
ame three types of vegetative organs of the angiosperms
1. roots
2. stems
3. leaves
angiosperms organized into two systems
1. shoot systems
2. root systems
shoot system
consists of stems and leaves in which photosythesis takes place
root systems
anchors and provides nutrients for the shoot system
morphology
structure of vegetative organs
and arrangement of component cells and tissues, or anatomy
name three root systems
1. taproots
2. fibrous root system
3. prop roots
taproots
single, large, deep-growing root, small side roots. The root may also function as food storage.

ex - sweet potato, carrot, beet
fibrous system
many thin roots of equal diameter. Have large surface area; cling to soil well

ex - green onion
prop roots
help support shoot
ex - corn
potato tuber
underground stem
runners
horizontal stems
name an adaptation of a plant that lives in the desert
desert plants have enlarged stems that store water
vascular plants - name tissue systems
1. vascular
2. dermal
3. ground

these systems extend throughout the plant body in a concentric arrangement
dermal tissue system
outer covering of the plant
vascular tissue system
conducts water and solutes thoughout the plant

the transport system

two parts, xylem and phloem
ground tissue system
carries out photosynthesis, stores photosynthetic products and helps support the plant
xylem
part of the vascular system

distributes water and minerals taken up by the roots to all parts of the plant. xylem can also function in storage and support
phloem
part of vascular system

transports carbohydrates from site of production (sources) to sites of utilization or storage (sinks)
how are plant cells unique?
1. chloroplasts
2. vacuoles
3. have cell walls with cellulose (differ in composition depending on function of cell)
describe cell walls in a plant
1. cell wall is outside the plasma membrane
2. cell walls can contain proteins and enzymes. chemical reactions can be important in cell expansion adn defense
3. except where waterproofed, cell walls are permeable to water and mineral ions
cell wall formation in plants
1. final step in mitosis
2. end of cytokenesis, two daughter cells separate by cell plate
3. each cell secrestes three polysaccharides - cellulose, hemicellulose, and pectin
4. cells expand and primary cell walls thin
5. expansion stops and second wall may be deposited
plasmodesmata
conect adjacent plant cells

cytoplasm-filled canals, traversed by a strand of ER

sometimes expand to allow macromolecules, including transcription factors and RNA, and viruses, to pass.

Doesn't require crossing a plasma membrane

allow direct communication between plant cells
what connects adjacent plant cells
plasomodesmata
plant cell types - name three
1. parenchyma cells
2. collenchyma cells
3. sclerenchyma cells
3a. fibers
3b. sclerids
4. xylem cells
4a. tracheids
4b. vessel elements
5. phloem cells
5a. sieve tube elements
parenchyma cells
numerous in young plants

have thin walls, large central vacuoles

photosynthetic cells in leaves are parenchyma cells with many chloroplasts

many store starch and lipids
starch is often stored in leucoplasts
leucoplasts
starch is often stored in these
collenchyma cells
have thick primary cell walls, usually elongate shape

provides support to leaf petioles, non-woody stems, and growing organs

tissue with collenchyma cells is flexible

ex strings of celery
sclerenchyma cells
thickened secondary walls

many die after secondary wall is laid down

provide STRONG support
sclerenchyma cells - name two types
1. fibers
2. sclerids
sclerenchyma cells - fibers
provides rigid support

ex - bark of tree
sclerenchyma cells - sclerids
packed together in a nuts shell or isolated clumps like the stones in cells of pears
xylem contains what types of elements, name element types and what do they do?
tracheary elements
1. tracheids
2. vessel elements

usually die before assuming their function
gymnosperm
meaning "naked seeds", after the unenclosed condition of their seeds (called ovules in their unfertilized state).
tracheids
gymnosperm have tracheid with pits in the secondary walls that allow materials to move freely
vessel elements
found in angiosperms

larger diameter than tracheid

end walls break down after death forming hollow tubes
seive tube elements
phloem transport cells

these are LIVING

cells meet end to end adn transport carbohydrates
meristem
undifferentiated plant tissue actively dividing (by mitosis)
early embriotic development - what two patterns contribute to basic body plan?
1. arrangement of cells and tissues along the main axis from root to shoot
2. concentric arrangement of tissue systems
modules (plant)
growing stem consists of these laid down one after another

each module is a node with attached leaves, the internode below, plus axillary buds at the base of that internode

think of a branch as a single module
how do branches differ from animal limbs
they form one after another, they differ in number of leaves, and number of subsequent branches
branches vs leaves - which is long lived, which is short?
branches - long-lived
leaves - short-lived (weeks to years)
root systems branching - describe
lateral roots may be semi-independent. as roots systems groww, roots may die and be replaced by new ones
all plants have a ____plat body
primary
which plants consist of primary plant body?
monocots
what has a secondary plant body and what does it consist of
trees and shrubs - consisting of wood and bark
secondary plant body
tissues are laid down as stems and roots thicken - grows throughout life
lateral meristems
cork cambium, vascular cambium
terminal bud
contains a shoot apical meristem
vascular cambium and cork cambium do what in what type of plants
thicken stem and root in woody plants
apical meristems
at tips of roots and stems, and in buds, give rise to the primary plant body
Shoot apical meristems
extend stems and branches
root apical ameristems
extends roots
name two types of apical meristems
1. Shoot apical meristems
2. root apical ameristems
what do apical meristems give rise to?
to a set of cylindrical primary meristems that produce three tissue systems
name three tissue types of primary meristems
protoderm
ground meristem
procambium
what does protoderm tissue turn into
dermal tissue system
what does ground meristem tissue turn into
ground tissue system
what does procambium tissue turn into
vascular tissue system
root cap
daughter cells become part of root cap - constantly erodes away
label
lateral meristems
produce secondary growth
vascular cambium
supplies cells of secondary xylem (toward the inside) which becomes wood; and secondary phloem (towards the outside) which becomes bark
cork cambium
produces waxy-walled cork cells - some of the cells become the bark
how do meristems build the plant body?
lateral meristems, vascular cambium, cork cambium
wood
secondary xylem
bark
everything external to the vascular cambium (periderm plus secondary phloem)
what represents primary and secondary growth?
a twig - has both - apical meristems are enclosed in bud

- only the buds are entirely primary tissues
buds of a plant are entirely what type of tissue?
primary tissue
label diagram
as tree grows - what happens to outermost layers?
outermost layers, including epidermis, dry and crack off
periderm
protective cells produced by cork cambium -

walls impregnated with suberin (waterproofing product)
meristems remain active for how long? - example of oldest plant
years or centeries

bristlecone pine - Pinus longaeva 4900 years old
root tissues:
protoderm
ground meristem
procambium
...give rise to
1. epidermis
2. cortex and endodermis
3. stele (vascular cylinder)
label
what does stele constist of
pericycle
xylem
phloem
pericycle
undifferentiated cells
1. give rise to lateral roots
2. gives riste to lateral meristems that thicken the root
3. cells have transport proteins that move nutrient ions into the xylem
label root tissues
label - what is this
eudicot stele
label - what is this?
monocot stele
label - what is this
monocot vascular bundles in stem
label - what is this?
eudicot stem vascular bundles
stems may have storage tissue in center called..
pith
cortex may also function as what - and what my they contain
they may function as storage and contain collenchyma cells
what constitutes the ground system
pith and cortex
secondary groth results from what?
from the lateral meristems in eudicots
what is vasuclar cambium initially?
initially a single layer of cells between primary xylem and phloem
how do secondary phloem and secondary xylem form and where?
when vascular cambium divide they form secondary phloem (toward outside) and secondary xylem (toward inside)
describe what (plant example) and why monocots do have thickened stems
ex - palms
- don't have vascular or cork cambiums - wide apical meristem produces wide stem, and dead leaf bases also contribute to the stem
how can you tell age of a tree in temerate region?
annual rings result from seasonal conditions
1. spring - water is plentiful - tracheids or vessel elements produced have large diameter
2. summer - less water - smaller diamter cells with thicker walls are produced
label
name experiment that describes how leaves relate to annual rings
ponderosa pine - needles connect with the xylem formed in teh year in which the needles were formed
why does center of tree look darker - what called
as tree grows in diameter, the xylem in the center becomes clogged with water insoluble substances -

heartwood
heartwood
center of tree - gets darker as tree gets older due to water insoluble substances
sapwood
xylem that actively conducts water
xylem that actively conducts water
sapwood
what are knots?
cross sections of branches
what are cross sections of branches
knots
how does leaf anatomy support photosynthesis
leaf anatomy is adapeted to carry out:
1. photosynthesis
2. exchange of O2 and CO2 with the environment
3. while limiting water losses
name two zones of cells to make up mesophyll - what type of cells
parenchyma - cell type

1. palisade mesophyll
2. spongy mesophyll

also include air for diffusion of gases
label - what is this
eudicot leaf cross section
vascular tissues form ___ in leaves
- where are they in plants
veins

- come within a few cell diamters of all the cells
how are mesophyl cells well supplied with water and mineral

- and why needed?
veins come within a few cell diamters of all the cells

products of photosyntheis can be conducted to the phloem
epidermal cells - describe function
nonphotosynthetic
- have waxy cuticle that is impermeable to water

- prevents water loss and diffusion of gas
what allows for gas exchange in a leaf
stomata
what are stomata controlled by
guard cells
if there is more solute on one side of the membrane - what does this mean in terms of water
less water if more solute and water will move in.
solute or osmotic potential
teh greater the solute concentration, the more negatie the solute potential
what is the driving force for osmosis to occur?
differeing concentrations of water
isotonic
water concentration same
hypertonic
high concentration of water
in cell - water moves out
hypotonic
conc of water is low in cell - water moves in
tugor pressure
water enters a cell due to its negative solute potential

entry of more water is resisted by an opposing tugor pressue
what happens if positive pressure potential is lost in a plant?
plant wilts
why would a plant wilt?
because it lost its positive pressure potential
pressure potential
over long dinstances in xylem and phloem, the flow of water and dissolved solutes is driven by this
wilted plant
neg water potential
=
neg solute potential
no pressue potential
well watered healthy plant
water potential =0
=
neg solute potential
equal pos pressure potential
bulk flow
movement of a solution due to difference in pressure potential
bulk flow in xylem is...
between regions of different negative pressure potential (tension)
bulk flow in phloem is...
between regions of different positive pressure potential (turgidity)
aquaporins
membrane channel proteins that water can pass through rapidly
abundance in plasma membrane and tonoplast depends on what
a cell's need to obtain or retain water
tonoplast
vacuole membrane
rate of water
direction of water
- which can be controlled
rate of water
how do mineral ions pass membranes?
transport proteins
concentration gradient
when molecules and ions move across membrane (from higher conc to lower) as permitted by membrane characteristics
active transport
conc of most ions in the soil solution is lower than that in the plant; uptate must be by this type of transport requiring energy
symport protein
together - protein in membrane that allows two molecules to pass through (ex H+ an Cl-)
how do water and minerals move into stele? name two methods
1. apoplast
2. symplast
apoplast
cell walls and intercellular spaces form a continuous meshwork that water can move through without crossing any membranes
symplast
water passes through cells via the plasmodesmata
label
what method must water use to enter stele
symplast
describe how leaves pull xylem sap upward
1. transpiration (water diffuses though stomata)
2.. water evaporates from mesophyll cells
3. tension pulls water from veins into mesophyll cells
4. tension ulls water upward into xylem of veins
5. tension pulls water up in xylem of the root and stem
6. water molecules form a cohesive water colum fromt eh roots to the leavs
7. water moves into the system by osmosis
8. water enters the root from soil by osmosis
guard cells - how do the open and close
in the presence of light, K+ and CL- are transported in, water follows due to neg water potential in the guard cell and opening cell

at night, K+ isn't transported, water potenial increases and water stops moving in - makes guard cell go limp and close
how are substances translocated in the phloem?
two steps in transloation require energy
1. transport of solutes from sources into sieve tubes (loading)
2. rmoval of solutes at sinks (unloading)

combination is pressure flow model
pressue flow model
combination of
1. transport of solutes from sources into sieve tubes (loading)
2. removal of solutes at sinks (unloading)
describe in 6 steps - the pressue flow model
1. transpiration pulls water to xylem vessels
2. source cells load sucrose into phloem sive tubes, reducing their water potential
3. so water is taken up from xylem vellels by osmosis - raising teh pressure potential in the sieve tubes
4. interal pressure differences dreive the sap down the sieve tube to sink cells
5. sucrose is unloaded into sink cells...
6. and water moves back to xylem vessels
where are sex organs in angiosperms
in the flowers
most angiosperms reproduce how?
sexually
sexual reproduction...define
produces new gene combinations and diverse phenotypes
asexual reproduction...define
produces clones of genetically identical individuals
examples of asexual propagation
strawberries
potatoes
bananas
navel oranges
examples of sexual reproduction (from seeds)
wheat
rice
corn
soybean
carpels
female sex organs
pistil
>1 carpel
stamens
male sex organs
petals
make up the corolla
corolla
made up with petals
sepals
make up calyx
calyx
small green "leaves" below petals... made of up sepals
name parts
gametophyte
haploid (n) phase that produces gametes
what does female gametophyte develop in?
megasporangium
what does male gametophyte develop in?
microsproangia
alternation of generations
multicellular diploid generations (2n) alternates with a multicellular haploid generation (n)
diploid (2n) generation

what is this called and what does it do?
sporophyte

it produces flowers
flowers produce what?
haploid spores (n)
spores develop into what?
into the haploid gametophyte (n) generation
megagametophytes develop in....
female
develop in megasporangia - embryo sacs
megasporangia are where what happens?
megagametophytes develop in

female

embryo sacs
microgametophytes develop in....
male
develop in microsporangia - pollen grains
microsporangia are where what happens?
microgametophytes develop in these

male

pollen grains
what is not required for transfer of pollen from anther to stigma
water is not required
name 2 types of transfer of pollen from anther to stigma
1. self pollination
2. cross pollination
self pollination
when plants fertilize themselves

ex - peas - occurs before flower opens
cross pollination
pollen is transfered to a different individual
mechanisms of pollination
1. wind (spring - allergies)
2. water (pollinate aquatic plants)
3. animals (insects, birds, bats)
self incompatibility

what are advantages?
what gene?
when plants reject their own pollen

- promotes outcrossing between different genotypes
- S gene
- many alleles
- if S allele in pollen matches either S allele in pistil, the pollen grain fails to grow
what happens after compatible sperm lands?

- where is sperm delivered?
germination occurs
- growth of a pollen tube

- sperm delivered to the ovule?
germination
when compatible sperm lands and pollen tube grows
pollen tube - where does it grow and go?
grows through the megasporangium and reaches the embryo sac
waht travels down pollen tube?

and what happens to it?
generative cell

- divides into two haploid sperm cells through miotic divsion
were do sperm enter after pollen tube?
both sperm cells enter a synergid, which degenerates and releases the sperm cells
what fertilization characteristic is that of angiosperms?
double fertilization

- two nuclear fusion events
what happens to two sperms?
- one fuses with the egg cell, forming a diploid zygote - divides mitotically to produce the sporophyte embryo
- other sperm fuses with the two polar nuclei forming a tripoid nucleus - divides by mitosis to form the nutritive endosperms (popcorn)
sporophyte embryo
one sperm fuses to form this
endosperm (popcorn)
one sperm fuses with polar nuclei forming a triploid nucleus (surrounds embryo - popcorn)

- accumulates starch, lipids, proteins
what physical characteristic is seen in the development of eudicots?
heart shaped embryo
what happens to cotyledons in seeds?
- some absorb nutrients from the endosperm and become much larger than the embryo (kidney bean)

- some remain thin, and use nutrients from the endosperm during germination (castor bean)
what do seeds do during development to ensure survival
in later stages of development, the seed loses water and becomes dormant
integuments - define and what does it develop into?
tissues surrounding the megasporangium - develop into the seed coat
carpel - what does it develop into?
becomes the wall of the fruit that surrounds the seed
fruit - describe what consists of
may contain only mature ovary and seeds or include other structures
name 4 ways fruits facilitate seed dispersal in 4 ways
1. water dispersal (coconuts)
2. animals (hide adn eat)
3. wind (parachute)
4. hitchhiking (on animals)
vegetative growth - what happens
apical meristems continuously produce leaves, stems, and axillary buds

- indeterminate growth
indeterminate growth
vegetative growth
determinate growth
if an apical meristem becomes an inflorescence meristem it produces bracts and new meristems in the angle between bract and stem
inflorescence meristems
give rise to floral meristems, bracts, and more inforescence meristems
floral meristems
gives rise to a flower
bract
modified usually reduced, leaflike structure
what gives rise to a flower
floral meristems
what gives rise to floral meristems, bracts, and more inflorescence meristems
inflorescence meristems
what triggers transition to the flowering state
environmental cues
name three types of flowering plants
1. annuals
2. biennials
3. perennials
annuals
complete life cycle in one year

- ex daisy, tomatoes
biennials
complete life cycle in two years

- parsnips, carrots
perennials
live for several to many years

- oak tree
control of flowering and other responses by length of day or night
photoperiodism
photoperiodism
control of flowering and othe responses by length of day or night
name example - experiment for photoperiodism
1920's mutant tobacco plants

- plants would not flower until day length was shorter than 14 hours

- critical day length
critical day length
length of day that is required for plant to flower
short day plants (SDPs)
flower when the day is shorter than a critical maximum.
- flower in late summer or fall

ex - mums, maryland tobacco
long day plants (LDPs)
flower when the day is longer than a critical maximum.
- flower in midsummer

ex. - spinach, clover
short-long-day plants
must experience short days flollowed by long days to flower
- spring (or just before mid-summer)

ex - white clover
long-short-day plants
plants must experience long days followed by short days
- bloom in fall
critical day length
length of day that is required for plant to flower
short day plants (SDPs)
flower when the day is shorter than a critical maximum.
- flower in late summer or fall

ex - mums, maryland tobacco
long day plants (LDPs)
flower when the day is longer than a critical maximum.
- flower in midsummer

ex. - spinach, clover
short-long-day plants
must experience short days flollowed by long days to flower
- spring (or just before mid-summer)

ex - white clover
long-short-day plants
plants must experience long days followed by short days
- bloom in fall
why is photoperiodism important
synchronizes flowering in the same species so that cross-pollination and successful reproduction is promoted
day-neutral plant
more common than photoperiodic plants - rely on other cues
what season do these flower?
1. short-long-day plants
2. long-day plants
3. short-day plants
4. long-short-day plants
1. spring
2. summer
3. fall
4. fall
do photoperiodic plants measure day or night? how do we know?
measure night
- experiment in 1920s with cockleburr (SDP) plants
day/night time varied while other held constant
- didn't matter day, but night had to be longer than 9 hours (critcal night length)
critical night length
hours of night that had to be experienced by plant to flower

- cockleburr - 9 hours of night

- hypothesis was wrong - "short day plants measure day length"
what happens if niterruption occured during dark period
nullified the effect of a long night
phytochrome
plant pigment
how could red light interrruptions be reversed
- far-red light - indicating a plant pigmen (phytochrome) was the receptor
circadian rhythms
all eukaryote cells, and some prokaryotes, have a "biological clock" that oscillates between two states every 12 hours
period
length of one cycle
amplitude
magnitude of change
all circadian rhythms
1. period is insensitive to temp, but lowering temp may reduce the amplitude
2. highly persistent - may persist for days even in darkness
3. can be entrained - chagned to match cycles other than 24 hour cycle
4. brif exposure to light can shift the peak of the cycle (phase shift)
phase shift
shift the peak of the cycle
give example of plant showing circadian rhythm
clover leaflets - folded at night, unfold during the day
night flowering
- flowers open when expected to get pollinated - night flowering plants pollinated by nocternal animals like bats and moths
circadian rythms must confer...
selective advantage

- research suggests that the advantage is the coordination of gene expression with the daily light-dark cycle

- ex. chlorophyll is not produced in teh dark
does the circadian clock help a plant interact with its environment?
- use experiment in answer
hypothesis " plants fix more carbon photo-sythetically when their circadian clock matches the environments light-dark cycle

- 2 mutant plants - one with long-cycle rythm, one with short-cycle

conclusion - "each mutant performed best under the cycle that corresponded to its geneticlaly determined circadian rhythm
phytochromes - what does this word mean (phyto? chrome?)
plant color
what interact with the plants biological clock
phytochromes and blue-light receptors interact with the plants biological clock
explain leaves measureing light/dark experiement
hypothesis "the leaves measure the dark period"

- masked one leaf - flowered
-masked one leaf and all plants flowered (when grafted together)
give examples of plants controlled by temp
wheat
rye

have two types of flowering behavior: annual and biannual

- spring wheat is annual
- winter wehat is biennial - planted in fall and flowers in sping
winter wheat
biennial - must be exposed to cold in its first year ot it will not flower the second
induction of flowering by low temperatures
vernalization

- it inhibits expression of a gene whose protein product represses other genes for flower development
asexual reproduction
produces a clone of progeny, genetically identical to the parent

- if well adapted to its environment, asexual reproduction can preserve and spread that successful genotype
advantages to asexual reproduction
1. not reliant on weather to bring pollen
2. not reliant on other plants nearby
3. happen quickly in good conditions
4. produces offspring more likely to mature
vegetative organs
stems, leaves, roots
what makes asexual vegetative reprodution possible
modification of vegetative organs

ex - strawberries - runer or stolons - develop into new plants
give example of vegetative reproduction
runners - strawberries
tip layers - blackberry
tubers - potatoes
rhizomes - bamboo
suckers - aspen tree
plantlets
bulb - onion
tip layers
branches that sag to the ground and a new plant grows from the branch tip
vernalization
induction of flowering by low temperatures

- it inhibits expression of a gene whose protein product represses other genes for flower development
rhizomes
horizontal underground stems that give rise to new shoots

ex. bamboo
bulbs
also known as corms
- short , vertical underground stems

- have fleshy, modified leaves for food storage - a large, underground bud. these can give rise to new plants (lilies, onions)
suckers
shoots produced by roots. many grasses and trees, such as aspens form interconnected stands of genetically identical individuals
where do asexual plants traditionally live

give example
unstable environments

- ex plants with stolons and rhizomes pioneers of sand dunes (beach grasses)

rapid reproduction allows them to survive shifting in sangs, and their roots help stabalize the dune
vegetative reproduction - disadvantages
example
all the same - so one virus or disease could wipe out all

- dutch elm disease (no genetic diversity to fight disease)
where is vegetative reproduction used

- how?
in agriculture

- stem cuttings inserted into soil will often grow into a new plant, especially if treated with auxin
grafting
root bearing plant is the stock

part grafted on is scion

vascular cambia of each must grow together so that water and minerals can be transported to the scion - usually closely related species
scion
part grafted
stock
root bearing plant
totipotent
undifferentiated cells
name ways totipotent cells are important
1. researchers are investigating ways to reproduce plants by tissue culture
2. culturing tiny bits of apical meristem can eliminate plant viruses
how have we used asexual plants to modify our produce
treatment with hormones can cause apical meristmes to produce millions of plants a year

produce virus-free, frost-resistant, increased nutritive value

ex. strawberries frost resistant- now plant wont die with early frost
plants ways to adapt to harsh environment
1. desert - self amputation
2. strangling fig in jungle
3. carnivorous plants - need nitrogen and can't get from soil